In the phenomenon of opioid tolerance, what causes opioid receptors to be recycled less?

The question relates to this popularisation video describing opioid tolerance.
In fact, reduced recycling of opioid receptors is just one of the processes involved in the development of opioid tolerance. The phenomenon of opioid tolerance is presented by the need to increase the dose of opioids in order to maintain the same level of analgesia. In a normal process, opioid receptors are rapidly desensitised following activation and then internalised in cells. Once the receptors have been internalised, they can be degraded (in structures known as lysosomes), recycled to the membrane and then resensitised. Although much remains to be discovered about this process, it is possible that the development of tolerance, as a function of time and the level of exposure of the receptors, implies that the degradation pathway is favoured (to the detriment of the recycling pathway). However, several studies have shown that in the central nervous system, the quantity of opioid receptors is not necessarily reduced despite the development of tolerance. In fact, the phenomenon of tolerance also involves a number of other adaptation mechanisms that result in analgesics having less effect.